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The dog (Canis lupus familiaris and Canis lupus dingo) is a domesticated form of the gray wolf, a member of the Canidae family of the order Carnivora. The term is used for both feral and pet varieties. The dog was the first animal to be domesticated, and has been the most widely kept working, hunting, and companion animal in human history. The word "dog" may also mean the male of a canine species, as opposed to the word "bitch" for the female of the species.
Dogs were domesticated from gray wolves about 15,000 years ago. They must have been very valuable to early human settlements, for they quickly became ubiquitous across world cultures. Dogs perform many roles for people, such as hunting, herding, pulling loads, protection, assisting police and military, companionship, and, more recently, aiding handicapped individuals. This impact on human society has given them the nickname "Man's best friend" in the western world. In 2001, there were estimated to be 400 million dogs in the world.
Over the 15,000 year span the dog had been domesticated, it diverged into only a handful of landraces, groups of similar animals whose morphology and behavior have been shaped by environmental factors and functional roles. Through selective breeding by humans, the dog has developed into hundreds of varied breeds, and shows more behavioral and morphological variation than any other land mammal. For example, height measured to the withers ranges from a few inches in the Chihuahua to a few feet in the Irish Wolfhound; color varies from white through grays (usually called "blue'") to black, and browns from light (tan) to dark ("red" or "chocolate") in a wide variation of patterns; coats can be short or long, coarse-haired to wool-like, straight, curly, or smooth. It is common for most breeds to shed this coat.
Etymology and related terminology
Dog is the common use term that refers to members of the subspecies Canis lupus familiaris (canis, "dog"; lupus, "wolf"; familiaris, "of a household" or "domestic"). The term can also be used to refer to a wider range of related species, such as the members of the genus Canis, or "true dogs", including the wolf, coyote, and jackals; or it can refer to the members of the subfamily Caninae, which would also include the African wild dog; or it can be used to refer to any member of the family Canidae, which would also include the foxes, bush dog, raccoon dog, and others. Some members of the family have "dog" in their common names, such as the raccoon dog and the African wild dog. A few animals have "dog" in their common names but are not canids, such as the prairie dog.
The English word dog comes from Middle English dogge, from Old English docga, a "powerful dog breed". The term may derive from Proto-Germanic *dukkōn, represented in Old English finger-docce ("finger-muscle"). The word also shows the familiar petname diminutive -ga also seen in frogga "frog", picga "pig", stagga "stag", wicga "beetle, worm", among others. Due to the archaic structure of the word, the term dog may ultimately derive from the earliest layer of Proto-Indo-European vocabulary, reflecting the role of the dog as the earliest domesticated animal.
In 14th-century England, hound (from Old English: hund) was the general word for all domestic canines, and dog referred to a subtype of hound, a group including the mastiff. It is believed this "dog" type of "hound" was so common it eventually became the prototype of the category “hound”. By the 16th century, dog had become the general word, and hound had begun to refer only to types used for hunting. Hound, cognate to German Hund, Dutch hond, common Scandinavian hund, and Icelandic hundur, is ultimately derived from the Proto-Indo-European *kwon- "dog", found in Welsh ci (plural cwn), Latin canis, Greek kýōn, Lithuanian šuõ.
In breeding circles, a male canine is referred to as a dog, while a female is called a bitch (Middle English bicche, from Old English bicce, ultimately from Old Norse bikkja). A group of offspring is a litter. The father of a litter is called the sire, and the mother is called the dam. Offspring are, in general, called pups or puppies, from French poupée, until they are about a year old. The process of birth is whelping, from the Old English word hwelp, (cf. German Welpe, Dutch welp, Swedish valp, Icelandic hvelpur).
The domestic dog was originally classified as Canis familiaris and Canis familiarus domesticus by Carolus Linnaeus in 1758, and was reclassified in 1993 as Canis lupus familiaris, a subspecies of the gray wolf Canis lupus, by the Smithsonian Institution and the American Society of Mammalogists. Overwhelming evidence from behavior, vocalizations, morphology, and molecular biology led to the contemporary scientific understanding that a single species, the gray wolf, is the common ancestor for all breeds of domestic dogs; however, the timeframe and mechanisms by which dogs diverged are controversial. Canis lupus familiaris is listed as the name for the taxon that is broadly used in the scientific community and recommended by ITIS; Canis familiaris, however, is a recognised synomym.
History and evolution
Domestic dogs inherited a complex social hierarchy and behaviors from their wolf ancestors. Dogs are pack hunters with a complex set of behaviors related to determining each dog's position in the social hierarchy, and they exhibit various postures and other means of nonverbal communication that reveal their states of mind. These sophisticated forms of social cognition and communication may account for their trainability, playfulness, and ability to fit into human households and social situations, and these attributes have given dogs a relationship with humans that has enabled them to become one of the most successful species on the planet today.
Although experts largely disagree over the details of dog domestication, it is agreed that human interaction played a significant role in shaping the subspecies. Shortly after domestication, dogs became ubiquitous in human populations, and spread throughout the world. Emigrants from Siberia likely crossed the Bering Strait with dogs in their company, and some experts suggest the use of sled dogs may have been critical to the success of the waves that entered North America roughly 12,000 years ago, although the earliest archaeological evidence of dog-like canids in North America dates from about 9,000 years ago. Dogs were an important part of life for the Athabascan population in North America, and were their only domesticated animal. Dogs also carried much of the load in the migration of the Apache and Navajo tribes 1,400 years ago. Use of dogs as pack animals in these cultures often persisted after the introduction of the horse to North America.
The current consensus among biologists and archaeologists is that the dating of first domestication is indeterminate. There is conclusive evidence dogs genetically diverged from their wolf ancestors at least 15,000 years ago, but some believe domestication to have occurred earlier. It is not known whether humans domesticated the wolf as such to initiate dog's divergence from its ancestors, or whether dog's evolutionary path had already taken a different course prior to domestication. For example, it is hypothesized that some wolves gathered around the campsites of paleolithic camps to scavenge refuse, and associated evolutionary pressure developed that favored those who were less frightened by, and keener in approaching, humans.
The bulk of the scientific evidence for the evolution of the domestic dog stems from archaeological findings and mitochondrial DNA studies. The divergence date of roughly 15,000 years ago is based in part on archaeological evidence that demonstrates the domestication of dogs occurred more than 15,000 years ago, and some genetic evidence indicates the domestication of dogs from their wolf ancestors began in the late Upper Paleolithic close to the Pleistocene/Holocene boundary, between 17,000 and 14,000 years ago. But there is a wide range of other, contradictory findings that make this issue controversial.
Archaeological evidence suggests the latest dogs could have diverged from wolves was roughly 15,000 years ago, although it is possible they diverged much earlier. In 2008, a team of international scientists released findings from an excavation at Goyet Cave in Belgium declaring a large, toothy canine existed 31,700 years ago and ate a diet of horse, musk ox and reindeer.
Prior to this Belgium discovery, the earliest dog fossils were two large skulls from Russia and a mandible from Germany dated from roughly 14,000 years ago. Remains of smaller dogs from Natufian cave deposits in the Middle East, including the earliest burial of a human being with a domestic dog, have been dated to around 10,000 to 12,000 years ago. There is a great deal of archaeological evidence for dogs throughout Europe and Asia around this period and through the next two thousand years (roughly 8,000 to 10,000 years ago), with fossils uncovered in Germany, the French Alps, and Iraq, and cave paintings in Turkey. The oldest remains of a domesticated dog in the Americas were found in Texas and have been dated to about 9,400 years ago.
A growing body of research indicates the companionship of a dog can enhance human physical health and psychological wellbeing. Dog and cat owners have been shown to have better mental and physical health than nonowners, making fewer visits to the doctor and being less likely to be on medication than nonowners. In one study, new pet owners reported a highly significant reduction in minor health problems during the first month following pet acquisition, and this effect was sustained in dog owners through to the end of the study. In addition, dog owners took considerably more physical exercise than cat owners and people without pets. The group without pets exhibited no statistically significant changes in health or behaviour. The results provide evidence that pet acquisition may have positive effects on human health and behaviour, and that for dog owners these effects are relatively long term. Pet ownership has also been associated with increased coronary artery disease survival, with dog owners being significantly less likely to die within one year of an acute myocardial infarction than those who did not own dogs.
The health benefits of dogs can result from contact with dogs, not just from dog ownership. For example, when in the presence of a pet dog, people show reductions in cardiovascular, behavioral, and psychological indicators of anxiety. The benefits of contact with a dog also include social support, as dogs are able to not only provide companionship and social support themselves, but also to act as facilitators of social interactions between humans. One study indicated that wheelchair users experience more positive social interactions with strangers when they are accompanied by a dog than when they are not.
The practice of using dogs and other animals as a part of therapy dates back to the late 18th century, when animals were introduced into mental institutions to help socialize patients with mental disorders. Animal-assisted intervention research has shown that animal-assisted therapy with a dog can increase a person with Alzheimer’s disease’s social behaviours, such as smiling and laughing. One study demonstrated that children with ADHD and conduct disorders who participated in an education program with dogs and other animals showed increased attendance, increased knowledge and skill objectives, and decreased antisocial and violent behavior compared to those who were not in an animal-assisted program.
Domestic dogs have been selectively bred for millennia for various behaviors, sensory capabilities, and physical attributes. Modern dog breeds show more variation in size, appearance, and behavior than any other domestic animal. Nevertheless, their morphology is based on that of their wild ancestors, gray wolves. Dogs are predators and scavengers, and like many other predatory mammals, the dog has powerful muscles, fused wrist bones, a cardiovascular system that supports both sprinting and endurance, and teeth for catching and tearing. Dogs are highly variable in height and weight. The smallest known adult dog was a Yorkshire Terrier, that stood only 6.3 centimetres (2.5 in) at the shoulder, 9.5 cm (3.7 in) in length along the head-and-body, and weighed only 113 grams (4.0 oz). The largest known dog was an English Mastiff which weighed 155.6 kilograms (343 lb) and was 250 cm (98 in) from the snout to the tail. The tallest dog is a Great Dane that stands 106.7 cm (42.0 in) at the shoulder.
Dogs are susceptible to various diseases, ailments, and poisons, some of which can affect humans. To defend against many common diseases, dogs are often vaccinated. Some breeds of dogs are prone to certain genetic ailments such as elbow or hip dysplasia, blindness, deafness, pulmonic stenosis, cleft palate, and trick knees. Two serious medical conditions particularly affecting dogs are pyometra, affecting unspayed females of all types and ages, and bloat, which affects the larger breeds or deep-chested dogs. Both of these are acute conditions, and can kill rapidly. Dogs are also susceptible to parasites such as fleas, ticks, and mites, as well as hookworm, tapeworm, roundworm, and heartworm.
Dogs are highly susceptible to theobromine poisoning, typically from ingestion of chocolate. Theobromine is toxic to dogs because, although the dog's metabolism is capable of breaking down the chemical, the process is so slow that even small amounts of chocolate can be fatal, especially dark chocolate.
Dogs are also vulnerable to some of the same health conditions as humans, including diabetes, dental and heart disease, epilepsy, cancer, hypothyroidism, and arthritis.
Despite their descent from wolves and classification as Carnivora, dogs are variously described in scholarly and other writings as carnivores or omnivores. Unlike obligate carnivores, such as the cat family with its shorter small intestine, dogs can adapt to a wide-ranging diet, and are not dependent on meat-specific protein nor a very high level of protein in order to fulfill their basic dietary requirements. Dogs will healthily digest a variety of foods, including vegetables and grains, and can consume a large proportion of these in their diet.
A number of common human foods and household ingestibles are toxic to dogs, including chocolate solids (theobromine poisoning), onion and garlic (thiosulphate, sulfoxide or disulfide poisoning), grapes and raisins, macadamia nuts, as well as various plants and other potentially ingested materials.
In domestic dogs, sexual maturity begins to happen around age six to twelve months for both males and females, although this can be delayed until up to two years old for some large breeds. This is the time at which female dogs will have their first estrous cycle. They will experience subsequent estrous cycles biannually, during which the body prepares for pregnancy. At the peak of the cycle, females will come into estrus, being mentally and physically receptive to copulation. Because the ova survive and are capable of being fertilized for a week after ovulation, it is possible for a female to mate with more than one male.
Dogs bear their litters roughly 56 to 72 days after fertilization, with an average of 63 days, although the length of gestation can vary. An average litter consists of about six puppies, though this number may vary widely based on the breed of dog. In general, toy dogs produce from one to four puppies in each litter, while much larger breeds may average as many as twelve.
Some dog breeds have acquired traits through selective breeding that interfere with reproduction. Male French Bulldogs, for instance, are incapable of mounting the female. For many dogs of this breed, the female must be artificially inseminated in order to reproduce.